Still have energy conservation behavior as vestigal leftover of the move to land.

Some desert dwellers have clubs on tails or thagomizers in place of a shark tail.  They often ironically come of as meek or bipolar to other landsharks because they can’t slap with their tails as a friendly warning

All landsharks, even ones living on the coast have a very recessed caudal keel.

Immune to sleep spells, or at least affected by them differently.

Have fully fallen into the orc hate as the two races compete often for protein sources in overlapping territories. Landsharks often depict orcs in art as resembling the demonic orcas of myth that menaced their ancestors.

It doesn’t help that the landsharkfolk are easy for orcs to not see as people and landshark livers selling well on the blackmarket.

Nor the fact that landsharks, in the logic of “its okay to steal if its from bad guys” often raid orc gardens.

Have okay eyesight, but consider vision one of the least important senses leading those with the resources to turn to magic alot for replicating other senses such as smell, sound and electromagnetic fields.This has led to them being commissioned by art enthusiasts and helped get them a source of income for their packs.

Speaking of sound, since their voices are so soft (real sharks can’t roar after all) all cultures put a lot of emphasis percussion instruments in conflict. Both in personal use and warfare. 

Travel alot so starting to turn towards using automatons to guard settlements when they move to another with the season.

More omnivorous than you’d expect. 

Culinary tradition more around making things easier to digest than about taste

Dorsal fins mostly vanished from evolution, but hey wear fake ones for ceremonial purposes.

(Not sure what domestic animals they would keep? What animals travel well?)

Landsharks in urban areas often find work as bouncers and bodyguards